Game, M., Smith, F.W. 2020. Loss of intermediate regions of perpendicular body axes contributed to miniaturization of tardigrades. Proceedings of the Royal Society B 287:1931 20201135.

Angelini, D.R., Smith, F.W. Theme and variation in the development of insect mouthparts. In: Insect Mouthparts: Form, Function, Development and Performance Krenn, Harald W. (Ed.), Springer, 2019.

Smith, F.W., Cumming, M., Goldstein, B. 2018. Analyses of nervous system patterning genes in the tardigrade Hypsibius exemplaris illuminate the evolution of panarthropod brains. EvoDevo 9:19.

Smith, F.W. 2018. Embryonic In Situ Hybridization for the Tardigrade Hypsibius exemplaris. Cold Spring Harbor Protocols 2018 (11), pdb. prot102350.

Smith, F.W., Gabriel, W. 2018. Embryonic Immunostaining for the Tardigrade Hypsibius exemplaris. Cold Spring Harbor Protocols 2018 (11), pdb. prot102343.

Smith, F.W., Bartels, P.J., Goldstein, B. 2017. A hypothesis for the composition of the tardigrade brain and its implications for panarthropod brain evolution. Integrative and Comparative Biology 57: 546-559.

Smith, F.W. and Goldstein, B. 2017. Segmentation in Tardigrada and diversification of segmental patterns in Panarthropoda. Arthropod Structure & Development 46: 328-340.

Smith, F.W., Boothby, T.C., Giovannini, I., Rebecchi, L., Jockusch, E.L., Goldstein, B. 2016. The compact body plan of tardigrades evolved by the loss of a large body region. Current Biology 26: 224-229.

Boothby, T.C., Tenlen, J.R., Smith, F.W., Wang, J.R., Patanella, K.E., Osborne Nishimura, E., Tintori, S.C., Li, Q., Jones, C.D., Yandell, M., Messina, D.N., Glassock, J., Goldstein, B. 2015. Evidence for extensive horizontal gene transfer from the draft genome of a tardigrade. Proceedings of the National Academy of Sciences 112: 15976-15981.

Jockusch, E.L., Smith, F.W., 2015. Hexapoda: Comparative aspects of later embryogenesis and metamorphosis. In: Evolutionary Developmental Biology of Invertebrates volume 5, Wanninger, Andreas (Ed.), Springer, 2015, pp. 111-208.

Smith, F.W., Jockusch, E.L., 2014. The Metameric pattern of Hypsibius dujardini (Eutardigrada) and its relationship to that of other panarthropods. Frontiers in Zoology 11: 1-16.

Smith, F.W., Jockusch, E.L., 2014. Hox genes require homothorax and extradenticle for body wall identity specification but not for appendage identity specification during metamorphosis of Tribolium castaneum. Developmental Biology 395: 182-197.

Smith, F.W., Angelini, D.R., Jockusch, E.L., 2014. A Functional genetic analysis in flour beetles (Tenebrionidae) reveals an antennal identity specification mechanism active during metamorphosis in Holometabola. Mechanisms of Development 132: 13-27.

Smith, F.W., Angelini, D.R., Gaudio, M.S., Jockusch, E.L., 2014. Metamorphic labral axis patterning in the beetle Tribolium castaneum requires multiple upstream, but few downstream, genes in the appendage patterning network. Evolution & Development 16: 78-91.

Angelini, D.R., Smith, F.W., Jockusch, E.L., 2012. Extent with modification: Leg patterning in the beetle Tribolium castaneum and the evolution of serial homologs. G3: Genes|Genomes|Genetics 2: 235-248.

Angelini, D.R., Smith, F.W., Aspiras, A.C., Kikuchi, M., Jockusch, E.L., 2012. Patterning of the adult mandibulate mouthparts in the red flour beetle, Tribolium castaneum. Genetics 190: 639-654.

Aspiras, A.C., Smith, F.W., Angelini, D.R. 2011. Sex-specific gene interactions in the patterning of insect genitalia. Developmental Biology 360: 369-380.